Bachelor of Arts, Trinity College, Dublin, 1956; Master of Science, Cornell Univercity, 1959; Doctor of Philosophy, Cornell Univercity, 1961; Master of Arts (honorary), Harvard University, 1971.
Senior scientist, Weizmann Institute Science, Rehovot, Israel., 1962-1965; assistant professor applied biology, division engineering and applied physics, Harvard University, 1966-1969; associate professor, Harvard University, 1969-1970; Gordon McKay professor applied biology, division applied science, Harvard University, since 1970. Member Executive Committee Tyler Ecology Prize.
Member American Association for the Advancement of Science, American Society Microbiology.
Married Muriel Harris, 1957. Children— Susan, Tricia, Dara.