Bachelor, Duke University, 1964. Doctor of Philosophy, Stanford University, 1968.
Postdoctoral fellow, Stanford University, Palo Alto, California, 1968-1971; postdoctoral fellow, Searle Research laboratories, High Wycombe, England, 1971-1973; postdoctoral fellow, Harvard University, Cambridge, Massachusetts, 1973-1974; assistant professor, U. Washington, Seattle, 1974-1978; associate professor, U. Washington, Seattle, 1978-1981; professor, U. Washington, Seattle, since 1981. Investigator Howard Hughes Medical Institute, Seattle, since 1976.
Fellow American Association for the Advancement of Science. Member National Academy of Sciences, American Academy Arts and Sciences.
Married Kathleen Lockwood, June 1963 (divorced 1970). 1 child, Carolyn Reeve Palmiter Engel.