Doctor of Veterinary Medicine, Iowa State University, 1959. Master of Science, Iowa State University, 1961. Doctor of Philosophy, Iowa State University, 1965.
Research associate Iowa State University, Ames, 1959—1961, assistant professor, 1962—1965, associate professor, 1966—1972, professor, since 1972, associate director, associate dean College Veterinary Medicine, 1990—1992, interim dean, 1992—1993, dean College Veterinary Medicine, 1993—2000, interim dean, dean College Agriculture, director Agricultural Experiment Station, 2000—2002, professor emeritus, 2004. Operating manager Veterinary Laboratory Inc., Remsen, 1961—1962. Postdoctoral fellow National Institute for Allergy and Infectious Diseases, Hamilton, Montana, 1965—1966.
Senior United States scientist Alexander von Humboldt Foundation, Bonn, Federal Republic Germany, 1975-1976. Chairman International Research Program on Comparative Mycoplasmology, 1982-1986. President Iowa State University Research Foundation, Ames, 1984-1986.
Howard Dunne memorial lecturer American Association Swine Practitioners, 1984. Member advisory board Secretary Agriculture, 1996-1999. Member strategic planning task force United States Department of Agriculture, 1997-1999, member safeguarding task force, 2001-2002, member implementation team, 2003-2004.
Board governors International Skating Union Foundation, 2004^.
Volunteer Union Station Homeless Shelter, Pasadena, California, 2007—2010. Member American College Veterinary Microbiologists (diplomate, vice chairman 1974-1975, secretary-treasurer 1977-1983), ACVM(President award of Excellence, 2007), American Society Microbiology (chairman division 1985-1986), International Organization Mycoplasmology (chair 1990-1992, Board Directors award 2002), American Veterinary Medical Association, American Association for the Advancement of Science, Osborn Research Club, Conference Research Workers in Animal Diseases (council member, president 1992, Dedicatee, 2007), Association American Veterinary Medical Colleges (president 1997-1998), Rotary Club(chair disaster), American Association for the Advancement of Science.
Married Karen Mae Paulsen, September 1, 1957. Children: Scott, Susan.