Bachelor in Computer Science and Mathematics, University of Southern Mississippi, 1973; Doctor of Philosophy in Biochemistry-Molecular Biology, Harvard University, 1982.
Research fellow department biochemistry and molecular biology, Harvard University, 1982-1985;
assistant professor genetics, Harvard Medical School, 1985-1990;
associate professor, Harvard Medical School, since 1990;
assistant molecular biologist, Massachusetts General Hospital, Boston, 1985-1990;
associate molecular biologist, Massachusetts General Hospital, Boston, 1990-1997;
associate director, Molecular Sciences Institute, Berkeley, California, since 1997. Consultant bacterial and yeast genetics Genetics Institute, Cambridge, Massachusetts, 1984-1993, science advisory board, since 1994. Consultant development biology, 1991-1993, advisorto science board, 1993-1994.
Ad hoc reviewer genetic molecular biology program National Science Foundation, since 1986. Ad hoc member microbiology and virology study American Cancer Society, 1989-1990, member molecular biology and genetics ad hoc study section, 1990-1992, member science advising committee on virology and molecular genetics, since 1993.