Biosystems Engineering, Swarthmore College, 1963; Doctor of Philosophy in Biophysics, University of Pennsylvania, 1968; postgraduate, Max-Planck-Institute for Psychiatry, Munich, 1968-1971.
Postdoctoral fellow, Max-Planck-Institute for Psychiatry, Munich, 1969-1971;
research scientist, Max Planck Institute for Behavioral Physiology, Seewiesen, Federal Republic of Germany., 1970-1977;
assistant professor biology, U. Oregon, Eugene, 1976-1980;
associate professor, U. Oregon, Eugene, 1980-1986;
director Institute Neurosci., U. Oregon, Eugene, 1985-1990;
professor biology, U. Oregon, Eugene, 1986-1990;
professor neuroscience, human biology and psychology, Stanford (California) University, since 1990;
Benjamin Scott Crocker professor human biology
head human biology program, Stanford (California) University, 1996. Director Public Health Svc. training grant, U. Oregon. Panel member and site visiting National Institute General Medical Science, National Institutes of Health.
Panel member National Institute Child Health and Development, National Institutes of Health, National Research Council, member VIS-C study section, 1988-1992. Fogarty senior vis.scientist National Institutes of Health, London, 1984-1985. Visiting professor anatomy, U. Colorado, 1983.
Visiting scientist Max-Planck-Institute for Behavioral Physiology, 1971-1973. Hilgard visiting professor Stanford University, 1987-1988.
Member New York Academy of Sciences, American Association for the Advancement of Science, Association Research Ophthamology and Vision, Animal Behavior Society, International Society Neurothology.
Married Anne Jones, May 25, 1969. Children: Lia, Anya.