Bachelor, Johns Hopkins University, 1964; Doctor of Philosophy, Case Western Reserve, 1968; Master of Arts, University of Pennsylvania, 1981.
Assistant professor biology, Johns Hopkins University, Baltimore, 1970-1974; associate professor biology, Johns Hopkins University, 1974-1980; professor biology, University of Pennsylvania, Philadelphia, 1980-1992; chairman biological Department, University Pennsylvania, Philadelphia, 1982-1987; professor-on-leave, University of Pennsylvania, Philadelphia, since 1992; chief science officer, Neose Pharmaceuticals, Inc., Horsham, PA., 1992-1994; chairman, Chief Executive Officer, Neose Pharmaceuticals, Horsham, since 1994. Member editorial board Quarterly Review Biology, Journal Molecular Recognition.
Member American Association for the Advancement of Science, Society for Development Biology, American Society for Cell Biology, Society for Complex Carbohydrates.
Married Hallam Hurt, July 14, 1981. Children: Adam, Rachel, Hallam.