Bachelor, Valparaiso (Indiana) University, 1964. Doctor of Philosophy, Northwestern University, 1969.
Postdoctoral fellow, Scripps Clinic and Research Foundation, La Jolla, California, 1969-1970; postdoctoral fellow, U. Rochester, New York, 1970-1972; assistant professor Medical School, U. Louisville, 1972-1978; associate professor Medical School, U. Louisville, 1978-1988; professor Medical School, U. Louisville, since 1988; vice chairman microbiology Department, University Louisville, 1989-1994; distong. teaching professor, since 1995.
Organizer Summer Institute Teachers, University Louisville, since 1988. Regional coordinator American Latvian Association, Washington, since 1991. Member American Association for the Advancement of Science, Wind River Conference on Prokaryotic Biology (executive council Estes Park, Colorado, since 1982), American Society for Microbiology, American Society for Cell Biology.
Married Patricia A. Williams, August 2, 1975. Children: Krista M., A. Krisjans.