Background
He was born in Darlington, County Durham and educated at school in Darlington and at Dulwich, gaining a Science Entrance Scholarship to Saint Thomas"s Hospital in 1890.
He was born in Darlington, County Durham and educated at school in Darlington and at Dulwich, gaining a Science Entrance Scholarship to Saint Thomas"s Hospital in 1890.
He later obtained degrees from both London University and Cambridge University. He became a house physician and then a Demonstrator at Saint Thomas"s in the Department of Physiology. He was also appointed to Lecturership in Pharmacology at Cambridge University, where he resided, going up to London to King"s College, London to deliver his lectures, where he held the post of Professor of Materia Medica and Pharmacology.
In 1919 he was appointed Reader in Pharmacology at Cambridge.
He was elected a Fellow of the Royal Society in 1911. His candidacy citation read: "Professor of Pharmacology.
Distinguished as a pharmacologist. Professor of Pharmacology, King"s College, London.
Assistant to the Downing Professor of Medicine, Cambridge.
Author of "A Manual of Pharmacology" (1906). Articles in Allbutt"s "System of Medicine," and Hale White"s "Pharmacology." "The Action of the Alkaloids from "Anhalonium Lewinii (Journ Physiol, volume xxv). "The Action of some New Preparations from "Cannabis Indica" (Brit Medical Journ, 1899).
"The Action of Poehl"s Spermine" (Journ Physiol, volume xxv).
"The Action of Indian Podophyllin" (Edin Medical Journ, 1900). "The Composition and Action of Orchitic Extracts" (Journ Physiol, volume xxvi).
"The Innervation of the Testis" (Proc Brit Association, 1901). "The Innervation of the Frog"s Stomach" (Journ Physiol, volume xxviii).
"Hypodermic Purgatives" (Brit Medical Journ, 1902).
"The Bronchial Muscles" (with T G Brodie, Journ Physiol, volume xxix). "Action of Drugs on Nerve cells and Nerve endings with special reference to Apocodeine" (Journ Physiol, volume xxx). "Vaso-motor Nerve to the Lung" (with T G Brodie, ibid, volume xxx).
"The Selective Action of Cocaine on Nerve Fibres" (ibid, volume xxxii).
"The Clioscribe" (with O Inchley, ibid, volume xxxii). "The Pathology of Asthma" (with T G Brodie, Transport Path Social, volume liv).
"The Biology-chemical Standardisation of Drugs" (Transport Pharm Confer, 1905). "The Biology-chemical Standardisation of Drugs" (with G South Haynes, 1905).
"Drug Fallacies" (Brit Medical Journ, 1906).
"The Mode of Action of Drugs" (Medical Magazine, 1907). "A Delicate Volume Recorder" (Journ Physiol, 1907). "The Gaseous Metabolism of the Mammalian Heart" (with J Barcroft, ibid, volume xxxv).
"The Action of Alcohol on the Circulation" (Journ Physiol, volume xxxv).
"The Action of Placental Extract" (Internat Congr Physiol, 1907)."
He was awarded Officer of the Most Excellent Order of the British Empire in 1919 for his contributions during World War I. He died in Whittlesford, Cambridgeshire in 1931.
Royal Society.