Bachelor, Yale University, 1957; Doctor of Medical Dentistry, Harvard University, 1961; postgraduate School Dentistry Medicine, Harvard University, 1961-1964; Doctor of Philosophy, Massachusetts Institute of Technology, 1967; Doctor Orthodontia (honorary), University Gotenburg, Sweden, 1988; Doctor of Dental Surgery (honorary), University Ghent, Belgium, 1992.
Research associate, Harvard University, 1961, 64; staff associate microbiologist, Forsyth Dental Center, Boston, 1964-1969; member of staff, Forsyth Dental Center, Boston, 1967-1969; associate professor dentistry, University of Michigan Dental School, Ann Arbor, 1969-1974; associate professor microbiology, University of Michigan Dental School, Ann Arbor, 1971-1974; professor microbiology and dentistry, University of Michigan Dental School, Ann Arbor, since 1974. Ad hoc reviewer National Institutes of Health.
Member American Society Microbiology, International Association Dental Research, (award in Dental Caries, 1994), American Association Dental Research (president 1987).
Married Rosemary Mercugliano, June 14, 1958. Children: Heidi, Christa, Dana.