AM, Harvard University, 1963; Doctor of Philosophy, Harvard University, 1967.
Research associate, Princeton (New Jersey) U., 1967-1969; research associate, University of Illinois, Urbana, 1969-1986; visiting assistant professor, University of Illinois, Urbana, 1987-1990; senior research scientist, University of Illinois, Urbana, since 1990.
Member American Association for the Advancement of Science, American Society for Biochemistry and Molecular Biology, American Association Microbiology.
Married Shau-Jin Chang, August 29, 1964. Children: Iris, Michael.