Bachelor of Science, Ohio State University, 1958; Doctor of Medicine, Tufts U., 1962.
Assistant professor, State University of New York, Buffalo, 1968-1973; staff physician, Tufts U. New England Medical Center, Boston, since 1973; professor School of Medicine, Tufts U., Boston, since 1980. Director National Institutes of Health Digestive Disease Basic Research Center at New England Medical Center/Tufts U.
Cons. National Institutes of Health, Washington. Captain Medical Corps, United States Army, 1966-1968. Fellow Infectious Disease Society American.
Member American Association Physicians, American Society for Clinical Investigation, American Gastroenterology Association, Royal Society Tropical Medicine.
Married Linda Plaut (divorced 1987). Children: Julie, John. Married Anetta Slone, June 18, 1989.