Bachelor of Arts in Biology, Brown University, 1956. Doctor of Philosophy, Rockefeller University, 1960.
From instructor to associate professor, Brandeis U., Waltham, Massachusetts, 1960-1976; professor biology, Brandeis U., Waltham, Massachusetts, since 1976; department chairman, Brandeis U., Waltham, Massachusetts, 1981-1984. Member of advisory panel for development biology National Science Foundation, 1974-1977, member of advisory panel for cell biology, 1992-1993.
(Book by Fulton, Chandler, Klein, Attila)
Member American Association for the Advancement of Science, Society for Development Biology, American Society Cell Biology, Marine Biological Laboratory, Society Protozoologists, American Iris Society, Phi Beta Kappa, Sigma Xi.
Married Margaretta Lyon, June 11, 1955. Children— Thomas, Margaretta, William. Married Elaine Lai, June 27, 1981.