Bachelor of Science in Zoology cum laude, University Louisville, 1972. Doctor of Philosophy in Evolutionary Biology, University Chicago, 1980.
Visiting assistant curator mammals, Field Museum Natural History, Chicago, 1981-1982; curatorial associate department geology, Field Museum Natural History, Chicago, 1982-1985; research associate department geology, Field Museum Natural History, Chicago, since 1986; visiting assistant professor, Purdue University, Hammond, Indiana, 1982-1985; visiting assistant professor, University of Illinois, Chicago, 1985-1987; associate professor, College College of DuPage, Glen Ellyn, Illinois, 1987-1988; visiting associate professor, Northwestern University, Evanston, Illinois, 1988-1989; associate director biological science, Northwestern University, Evanston, Illinois, since 1989.
President Kennicott Club, Chicago, 1983-1985, 88-90, vice president, 1986-1988. Member Chicago Rainforest Action Group, since 1988, president, since 1991. Member American Association for the Advancement of Science, American Society Mammalogists, Paleontological Society, Organization Tropical Studies, Sigma Xi.
Married Sharon Kay Walker, August 3, 1985.