Bachelor of Science, University Minnesota, 1951; Doctor of Philosophy, University Minnesota, 1960; Master of Science, Colorado State University, 1953.
Research biologist, Minnesota Department Natural Resources, St. Paul, 1954-1956; ecologist, Minnesota Museum Natural History, Minneapolis, 1956-1961; lecturer to professor, head department ecology and behavioral biology, director Cedar Creek Natural History Area, University of Minnesota, Minneapolis, since 1961; vice president, State Bank Gibbon, Minnesota, 1966-1983; president, State Bank Gibbon, Minnesota, since 1983; also director, State Bank Gibbon. Member behavioral science training committee National Institutes of Health, 1968-1973.
Fellow American Association for the Advancement of Science, Aberdeen University (Scotland) (honorary). Member Ecological Society of America, American Institute Biological Science, Wildlife Society, American Society Mammalogists, New York Academy of Sciences, Minnesota Academy of Sciences, Explorers Club, Sigma Xi.
Married Joyce Jeanette Lund, September 12, 1959. Children: Hans, Peter.