Bachelor of Science with honors, University Manchester, 1961. Master of Science, University Manchester, 1962. Doctor of Philosophy, University Manchester, 1964.
Assistant lecturer University Sheffield, 1964-1965. Research associate University Washington, 1965-1967. Assistant lecturer University Glasgow, 1967-1968, lecturer, 1968-1978, senior lecturer, 1978—2004, honorary senior research fellow, since 2004.
Member Institute Biology, British Electrophoresis Society (secretary, 1989-1992, president 1994-1997), International Association for Dental Research (councillor 1995-1998), Association of Basic Science Teachers in Dentistry (secretary-treasurer since 1988).
Married John Gervais Beeley, September 15, 1965. 1 child, James Malcolm.