Bachelor of Arts, University of Pennsylvania, 1962, Doctor of Philosophy, 1966. Postdoctoral fellow Weizmann Institute Science, Rehovoth, Israel, 1966-1967. Postdoctoral associate Institute for Cancer Research, Philadelphia, 1968-1970, research associate, 1970-1972, assistant member, 1972-1977, associate member, 1977-1978.
Assistant professor biophysics University of Pennsylvania, Philadelphia, 1974-1978.
Associate professor of chemistry University of California, Berkeley, 1978-1982, professor, 1982, professor molecular and cell biology, 1993. Member ad hoc biochemistry and physical biochemistry study sections National Institutes of Health, 1977-1984, physical biochemistry study section, 1984-1988.
Postdoctoral fellow Weizmann Institute of Science, Rehovoth, Israel, 1966—1967. Postdoctoral associate Institute Cancer Research, Philadelphia, 1968—1970, research associate, 1970—1972, assistant member, 1972—1977, associate member, 1977—1978. Assistant professor biophysics University Pennsylvania, 1974—1978.
Associate professor chemistry University California, Berkeley, 1978—1982, professor, since 1982, Miller professor, 1992, 2003—2004, professor molecular and cell biology, since 1993, chair chemical department, 2000—2003, Joel Hildebrand chair, 2002—2003, chancellor's professor, since 2009. Member ad hoc biochemistry and physical biochemistry study sections National Institutes of Health, 1977—1984, physical biochemistry study section, 1984—1988.
Member editorial board Journal Biological Chemistry, 1979-1984, Biofactors, 1991, European Journal Biochemistry, 1991-1995, Biochemistry, 1993, Annual Review of Biochemistry, 1996. Fellow National Science Foundation, 1964, National Institutes of Health, 1964-1966. Member National Academy of Sciences, American Chemical Society (executive coun.biol. division 1982-1985, chairman nominating committee 1987-1988, program chair 1991-1992, Repligen award 1994), American Academy Arts and Sciences, American Society Biochemistry and Molecular Biology (membership committee 1984-1986, public affairs committee 1987-1994, program committee 1995, president-elect 1997, president 1998), Sigma Xi.
Married Norman R. Klinman, July 3, 1963 (divorced 1978). Children: Andrew, Douglas.