Doctor of Philosophy, University Tübingen, Tübingen, Germany, 1960., University Hamburg and Tübingen. Professor, University Tübingen, Tübingen, Germany, 1979.
Postgraduate, University Michigan, 1963.
Assistant department of botany University Tübingen, Germany, 1963-1966. Department of biology Princeton (New Jersey) University, 1966-1967. Habilitation, privatdozent University Tübingen, 1972, professor, 1979-2001.
Sabbatical leave Department of Zoology, Auckland, New Zealand, 1984, Department of Physics, Trondheim, Norway.
Married Sigrun Margund Hörr, July 25, 1938. Children: Dirk Friedrich, Lerke Hedwig.