Bachelor of Science, Iowa State University, 1964. Master of Science, Cornell University, 1966. Doctor of Philosophy in Soil Microbiology, Cornell University, 1968.
From assistant professor to professor, Michigan State University, 1968-1978;
distinguished professor, Michigan State University, since 1991;
director science and technical center microbial ecology, National Science Foundation, since 1988. Visiting associate professor of University Georgia, 1974-1975. Consultant National Science Foundation, 1974-1977.
Visiting professor University of California Berkeley, 1981-1982. Member biotechnology science advising committee Environmental Protection Agency, 1986-1989, chair science advisory county Grade Point Average, 1988-1990.
Member American Association for the Advancement of Science, American Society Agronomy (Soil Science award 1990), International Institute Biotechnology, American Society Microbiology (award in applied and environmental microbiology 1992), Soil Science Society of America, Ecological Society of America, International Society Soil Science (chair soil biology division).
Married, 1965; 3 children.