Bachelor, Whitman College, Walla Walla, Washington, 1969. Master of Science, University Puget Sound, Tacoma, 1972. Doctor of Philosophy, University Washington, Seattle, 1977.
Research assistant College Forest Resources, University Washington, Seattle, 1972—1976. Senior conservation biologist Alaska Department Fish & Game, Anchorage, 1990—1997, research wildlife biologist Juneau, 1990—1997, game biologist, 1996—1997. Executive director Audubon Alaska, 1997—1999, senior scientist, since 1999.
Associate professor, wildlife biology University Alaska Fairbanks, since 1990.
Vice president Alaska Sea Life Center, Seward, 2001—2007. Member of Wildlife Society (Alaska chapter president 1982, fellowship 2006).
Married Mary Beth Schoen, September 6, 1970. Children: Erik R., Sarah K.