Education
Willadsen graduated from the Royal Veterinary College of Copenhagen (1969), and received a Doctor of Philosophy in reproductive physiology from there (1973).
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Steen Willadsen
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In 1984, at the British Agricultural Research Council"s Institute of Animal Physiology, Cambridge, he successfully used cells from early embryos to clone sheep by nuclear transfer. The procedure he developed was essentially the one used a decade later by Wilmut et al. to produce Dolly, the sheep, although in the latter case, nuclei from a mature sheep, id est (that is) not from sheep embryos, were used. Prior to the nuclear transfer experiments, Willadsen had developed methods for freezing sheep and cow embryos, and embryo manipulation methods for producing genetically identical animals, primarily identical twins in sheep, cattle, pigs, and horses, and for producing mammalian chimaeras, including interspecies chimaeras.
